Abstract The earliest history of Archosauriformes is mainly represented by members of Proterosuchidae and Erythrosuchidae, which are known worldwide from latest Permian to Middle Triassic beds. These two groups were historically combined within ‘Proterosuchia’, with approximately 30 nominal species. Two morphotypes have been recognized among proterosuchians: proterosuchids with a generally more sprawling gait and elongated and low skulls with an overhanging premaxilla, and the more heavily built erythrosuchids, with a probably less sprawling gait and large, presumably hypercarnivorous, skulls. The systematics of ‘Proterosuchia’ was relatively chaotic throughout most of the twentieth century, but currently there exists consensus regarding the non-monophyly of proterosuchians and their phylogenetic position outside all other archosauriforms. In contrast, the delimitation and taxonomic content of Proterosuchidae and Erythrosuchidae remain unstable. Few studies of proterosuchian palaeobiology have been carried out. Current lines of evidence favour a predominantly terrestrial lifestyle for proterosuchians. Limb bone histology indicates rapid continuous growth rates in Proterosuchus and Erythrosuchus before reaching sexual maturity. A better knowledge of proterosuchian anatomy, systematics, evolution and ecology is important for advancing understanding of the origin and early radiation of Archosauriformes and the patterns of biotic recovery following the Permo-Triassic mass extinction event. There remains much research to be carried out in proterosuchian palaeobiology.

Abstract Euparkeria capensis has long been considered an archetype for the ancestral archosaur morphology, and has been placed just outside of crown Archosauria by nearly all cladistic analyses. Six species are currently considered to be putative members of a clade Euparkeriidae, and have been collected from Olenekian- or Anisian-aged deposits in South Africa ( Euparkeria capensis – the only definitive member of the group), China ( Halazhaisuchus qiaoensis , Wangisuchus tzeyii , ‘ Turfanosuchus ’ shageduensis ), Russia ( Dorosuchus neoetus ) and Poland ( Osmolskina czatkowicensis ). Four other species ( Turfanosuchus dabanensis , Xilousuchus sapingensis , Platyognathus hsui , Dongusia colorata ) were historically assigned to Euparkeriidae, but have been removed by recent work. Recent authors deemed Osmolskina czatkowicensis and Dorosuchus neoetus to be the most likely taxa to form a euparkeriid clade with Euparkeria capensis , but Osmolskina czatkowicensis and Euparkeria capensis were not found as sister taxa by the only cladistic analysis to have tested euparkeriid monophyly. Euparkeria capensis was small (<1 m), insectivorous or carnivorous, probably had vision adapted to low-light conditions and a semi-erect crocodile-like stance, and may have been facultatively bipedal. Bone histology demonstrates that Euparkeria capensis had a slow growth rate, which has been suggested to have been an adaptation to relatively stable environmental conditions.

Abstract Phytosauria is a nearly cosmopolitan clade of large, quadrupedal, carnivorous archosauriforms. They are known unambiguously from Late Triassic deposits, although the clade’s ghost lineage extends at least to the late Early Triassic. Their nares are uniquely located close to the orbits rather than anteriorly in the rostrum as in modern crocodylians, and the rostrum is formed by elongated premaxillae bearing many teeth. Phytosaurs have roughly triangular, ornamented paramedian osteoderms, rounder appendicular osteoderms, and a unique ‘gular shield’ assembled from multiple, irregular osteoderms under the throat. Phytosaurs are reconstructed as semi-aquatic because of their general similarity to modern crocodylians and common preservation in fluvial and shallow-marine deposits. Currently, over thirty species are recognized. New specimens continue to be collected, some representing new taxa. The taxonomic status of other named taxa is uncertain and requires re-investigation. Since their discovery, phytosaurs have been used as biostratigraphic and biochronological index taxa for correlating Late Triassic sediments worldwide. Recent systematic and taxonomic revisions cast doubt on some of those correlations. Our understanding of the evolution of Phytosauria is far from complete. With reevaluation of well-known specimens, rigorous and comparative morphological descriptions, and robust phylogenetic hypotheses of ingroup relationships, studies of phytosaurs can address larger palaeobiological questions.

Abstract We present the first comprehensive description of Prorotodactylus and Rotodactylus dinosauromorph tracks from the Early and Middle Triassic of the Holy Cross Mountains, Poland. We describe and comprehensively figure tracks that have been mentioned briefly in previous accounts as well as new, recently discovered material, and analyse the variation and stratigraphic distribution of these specimens. Tracks have been recorded from four sites – Koszary, Stryczowice, Wióry and Baranów – which span the early Olenekian–early Anisian ( c. 250–246 Ma). These tracks therefore represent an ichnological record of the evolutionary succession of early dinosauromorphs during the earliest part of their evolutionary history. Recognized track types include cf. Prorotodactylus isp . , Prorotodactylus isp., Prorotodactylus mirus , Rotodactylus cursorius , Rotodactylus isp. and cf. Rotodactylus isp. At least three distinct Early and early Middle Triassic early dinosauromorph ichnofaunas can be recognized. The oldest, which is early Olenekian in age, is characterized by the presence of Prorotodactylus isp., cf. Prorotodactylus isp . and non-archosaurian archosauromorph or archosaur tracks (e.g. Synaptichnium isp., Protochirotherium isp.), recorded at the Stryczowice and Koszary sites. The following assemblage, recorded at the late Olenekian Wióry site, displays the highest ichnodiversity of dinosauromorphs, with four track types present ( Prorotodactylus isp., Prorotodactylus mirus , Rotodactylus cursorius and cf. Rotodactylus isp.). The youngest site, Baranów, includes Rotodactylus isp., as well as other larger dinosauromorph tracks. The first body fossil evidence of dinosauromorphs is a few million years younger than the youngest Polish tracks, so Prorotodactylus and Rotodactylus tracks currently provide the oldest record of dinosauromorph morphology, biology and evolution.

Abstract Mesozoic terrestrial vertebrates gave rise to sea-going forms independently among the ichthyosaurs, sauropterygians, thalattosaurs, crocodyliforms, turtles, squamates, and other lineages. Many passed through a shallow marine phase before becoming adapted for open ocean life. This allows quantitative testing of factors affecting our view of the diversity of ancient organisms inhabiting different oceanic environments. We implemented tests of correlation using generalized difference transformed data, and multiple regression models. These indicate that shallow marine diversity was driven by changes in the extent of flooded continental area and more weakly influenced by uneven fossil sampling. This is congruent with studies of shallow marine invertebrate diversity and suggests that ‘common cause’ effects are influential in the shallow marine realm. In contrast, our view of open ocean tetrapod diversity is strongly distorted by temporal heterogeneity in fossil record sampling, and has little relationship with continental flooding. Adaptation to open ocean life allowed plesiosaurs, ichthyosaurs and sea turtles to ‘escape’ from periodic extinctions driven by major marine regressions, which affected shallow marine taxa in the Late Triassic and over the Jurassic–Cretaceous boundary. Open ocean taxa declined in advance of the end-Cretaceous extinction. Shallow marine taxa continued diversifying in the terminal stages due to increasing sea-level. Supplementary material: The data series and full analytical results are available at http://www.geolsoc.org.uk/SUP18486