Questions of biotic and environmental change during deposition of the Upper Maastrichtian Hell Creek Formation require a robust and replicable system for intra-formational correlation of fossil localities. In this paper, we present a carbon isotope chemostratigraphic curve based on terrestrial organic carbon. Data were taken from a complete measured section spanning the full 93 m of the Hell Creek Formation at our study site. Sedimentary beds were described at the centimeter scale, and samples for carbon isotope analysis were taken at ~10 cm intervals. Each sedimentary bed was analyzed in thin section, and grain-size data were assembled based on petrographic point counts. The well-documented Cretaceous-Paleogene boundary negative carbon isotope excursion, six negative carbon isotope excursions, and four tentative positive carbon isotope excursions provide chronostratigraphic tie points within the Hell Creek Formation. We used this curve to precisely correlate 12 additional fossil-bearing localities from throughout the Hell Creek Formation across its type area. These correlations revealed significant local variation in sediment accumulation rates, confirming that simple stratigraphic position relative to the diachronous base and top of the Hell Creek Formation introduces significant error in correlation.

The Hell Creek Formation in eastern Montana has yielded well-preserved leaf megafossil localities that provide insight into the vegetation and climate of the latest Cretaceous. Among the most basal, the PDM locality (UCMP [University of California Museum of Paleontology] PB99057 = MOR [Museum of the Rockies] HC-278) occurs in channel sandstones ~10 m above the underlying Fox Hills Formation. The locality represents a fluvial/estuarine environment. Leaf megafossil impressions were preserved on clay drapes within the channel. Angiosperms dominated the flora (13 of 17 morphotypes). Dryophyllum subfalcatum and “ Vitis ” stantoni , two common morphospecies in the Hell Creek Formation, are well represented. Gymnosperms including Metasequoia , Glyptostrobus , Cupressinocladus , and Ginkgo are rare; ferns and cycadophytes are absent. Univariate leaf-margin analysis produced mean annual temperature (MAT) estimates of ~7–11 °C (5–14 °C including overlapping estimation errors). The Climate Leaf Analysis Multivariate Program (CLAMP) produced a MAT value of 11–12 °C ± 2 °C. Leaf area analysis produced mean annual precipitation (MAP) estimates of 197 cm (+152/−86 cm) and 191 cm (+161/−87 cm), while CLAMP produced a growing season precipitation estimate of 82–90 ± 48 cm. The wetter MAP values are consistent with paleosols near the base of the formation, which lack paleosol carbonate. CLAMP results further suggest seasonality in both temperature and precipitation. Some PDM morphotypes are familiar from the Hell Creek I floral zone of North Dakota, and several are not, suggesting greater spatial and/or temporal heterogeneity in the Hell Creek Formation flora than has been previously appreciated.

Many workers consider the Cretaceous-Paleogene extinction the archetypal catastrophic pulse event caused solely by the Chicxulub bolide impact. However, based on a global scale analysis of marine animals, the Cretaceous-Paleogene boundary is a candidate for an extinction enhanced by the coincidence of press and pulse disturbances. We make a preliminary test of key predictions of the press-pulse hypothesis using palynological data. We document a local palynological extinction of 21% at the Cretaceous-Paleogene boundary, which is consistent with extinction rates of 15% to 30% at other localities in the Hell Creek type area and throughout North America. We also find a decline in the number of dicot angiosperm pollen taxa between −3.5 m and −2.5 m below the boundary. We document a low-palynospecies-richness interval between −1.4 m and −1.0 m that includes extirpation, but not extinction, of some palynospecies. These changes in species richness are not correlated with changes in depositional style or pollen preservation, indicating that they may represent a biological rather than entirely taphonomic signal. Review and reanalysis of previously published data from other localities in the western interior of North America suggest similar declines in species richness within approximately the same stratigraphic interval. However, many of the species absent during the low-species-richness interval reappeared before the Cretaceous-Paleogene boundary, suggesting changes in community structure and composition before the terminal Cretaceous event—a key prediction of the press-pulse hypothesis—rather than gradual extinction in the latest Cretaceous.

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