Abstract Ordovician studies in Iran have shown significant progress since the beginning of the century. A number of individual faunas have been documented and a biostratigraphical framework based on conodonts, chitinozoans, acritarchs and trilobites developed. Correlation of Ordovician successions with the International Chronostratigraphic Chart has been significantly improved, and the position of the series and stage boundaries can be recognized with greater precision. While geographical proximity to temperate latitude Gondwana is apparent for most Iranian terranes, biogeographical links of Alborz and Kopet-Dagh with South China prevailed through the Early–Middle Ordovician. In Pakistan, Ordovician deposits have a restricted distribution in the Karakorum block (Chitral). Here they are represented by the Yarkhun and Vidiakot formations with Floian–Darriwilian acritarchs, chitinozoans and early Darriwilian conodonts. In Peshawar District of the North-West Frontier Province, an Early–Middle Ordovician age is likely for the Misri Banda Quartzite with Cruziana rugosa trace fossils. It is overlain conformably by carbonates of the Panjpir Formation, which has an inferred Middle Ordovician–Silurian age. Presently available information on the Ordovician of Afghanistan is mostly based on reconnaissance studies performed almost half a century ago, and a few monographed Early and Late Ordovician faunas.

Abstract The region of Kyrgyzstan, Tajikistan and Uzbekistan includes five first-order tectonic units with an Early Paleozsoic sedimentary record, comprising North Tien Shan, Karatau Naryn, Turkestan–Alai, Zeravshan–Hissar and the Central Pamirs. Available palaeobiogeographical and palaeomagnetic data suggest that these were widely dispersed in the Ordovician. North Tien Shan, Karatau Naryn, Turkestan–Alai were separate microcontinents located in the low southern latitudes throughout the Ordovician in relative proximity to the western Gondwana margin. Zeravshan–Hissar and the Central Pamirs were also parts of the Gondwana supercontinent but were located in temperate latitudes. The geological literature on the Ordovician of the region is assessed to provide an updated palaeontological record, outline of lithostratigraphy and biostratigraphic correlation based on the International Chronostratigraphic Chart. The Ordovician biostratigraphy of Central Asia is mainly graptolite-based; however, that record is discontinuous, and the absence of detailed faunal logs and lack of monographic studies causes difficulty in precisely locating system and stage boundaries. Although an extensive faunal record has been documented, often it is based on preliminary taxonomical identifications which are not reliable for high-resolution biostratigraphy and tracing biodiversity patterns.

Abstract The biogeographical patterns shown by Ordovician linguliform and craniiform brachiopods are greatly influenced by their dominance in low-diversity associations in marginal environments. This is particularly evident in the Early Ordovician, when linguliform-dominated dysaerobic assemblages are widely distributed along the deep shelves of Gondwana, the Kazakhstanian terranes and in Baltica. By the Darriwilian, micromorphic linguliforms are characteristic components of the pantropical climatic-controlled faunas of Laurentia, Cuyania and Kazakhstanian terranes, which – in spite of separation by extensive oceans – retain a distinct similarity. Analysis of craniiform biogeographical distribution is impeded significantly by the poor state of craniide taxonomy and lack of reliable data from most regions. However, in general their biogeographical dispersion is similar to other groups of the Palaeozoic Evolutionary Fauna. Unlike the linguliforms, which are important members of the Cambrian Evolutionary Fauna, there is no convincing Cambrian craniiform record; they may have evolved and dispersed from Gondwana and associated microcontinents and island arcs. The earliest well-established record is from the late Tremadocian of temperate to high-latitude peri-Gondwana. During most of the Ordovician, they have a peri-Iapetus distribution. They are very rare or absent in tropical Gondwana, South China and Kazakhstanian terranes and are not yet documented from Siberia. The trimerellides probably evolved in tropical peri-Gondwanan island arc settings. Their dispersion and major features of biogeography mirror those of atrypides. Gold Open Access: This article is published under the terms of the http://creativecommons.org/licenses/by/3.0/ .

Abstract Palaeobiogeographical data on Cambrian trilobites obtained during the twentieth century are combined in this paper to evaluate palaeoceanographic links through c. 30 myr, once these arthropods biomineralized. Worldwide major tectonostratigraphic units are characterized at series intervals of Cambrian time and datasets of trilobite genera (629 for Cambrian Series 2, 965 for Cambrian Series 3, and 866 for the Furongian Series) are analysed using parsimony analysis of endemicity. Special attention is given to the biogeographical observations made in microcontinents and exotic terranes. The same is done for platform-basinal transects of well-known continental margins. The parsimony analysis of endemicity analysis resulted in distinct palaeogeographical area groupings among the tectonostratigraphic units. With these groupings, several palaeobiogeographical units are distinguished, which do not necessarily fit the previously proposed biogeographical realms and provinces. Their development and spatial distributions are broadly controlled by Cambrian palaeoclimates, palaeogeographical conditions (e.g. carbonate productivity and anoxic conditions) and ocean current circulation. Supplementary material: Global dataset of Cambrian Epoch 2 (A), Cambrian Epoch 3 (B) and the Furongian Epoch (C) trilobite genera are provided at: http://www.geolsoc.org.uk/SUP18669

Abstract Two separate tectonic blocks in the southwestern segment of the Kazakhstanian orogen, the Chu–Ili terrane and the Karatau–Naryn terrane (with particular attention to Malyi Karatau), are selected to illustrate their geological history and major biogeographical signatures from the Cambrian to the early Silurian. Mid- to Late Ordovician brachiopod and trilobite faunas of Chu–Ili show increased endemicity of shallow shelf assemblages, whereas distinct links to equatorial (‘east’) peri-Gondwanan are more evident in trilobite assemblages of the outer shelf. In the Late Ordovician, strong biogeographical affinities to equatorial Gondwanan faunas became firmly established and they are also traceable into the Silurian. Early Cambrian faunas of Malyi Karatau show remarkable similarity to those of South China. From the Middle Cambrian this region evolved as an isolated carbonate seamount, but until the Early Ordovician links to South China faunas were still evident. Benthic faunas from both regions show weak links to contemporaneous faunas of Baltica and little in common with Cambrian and Ordovician faunas of the Siberian craton. This suggests their location in low southern latitudes, in relative proximity to East Gondwana, which places some constraints on plate-tectonic reconstructions in relation to the southern cluster of Kazakhstanian terranes, including Karatau–Naryn, North Tien Shan and Chu–Ili.

Abstract Brachiopod-dominated palaeocommunities incorporating a structure typical of faunal groups within the Palaeozoic Evolutionary Fauna were already present in North and East Gondwana and associated terranes as early as the mid-Cambrian, confined exclusively to shallow marine, inshore environments. The late Cambrian and Tremadoc record of these faunas is incomplete, because of pronounced global sea-level lowstand and subsequent break-up and destruction of the Cambrian Gondwanan margin. It is likely, however, that those groups later forming the core of the Palaeozoic Evolutionary Fauna evolved originally in shallow-water environments of low-latitude peri-Gondwana, and dispersed widely when favourable ecological conditions developed. Conspicuous sea-level rise through the early to mid-Arenig provided newly available habitats in the expanding epeiric seas, where the new faunas evolved and diversified by the mid-Ordovician, when rapid drift separated the early Palaeozoic continents. Relatively short-lived precursor and transitional brachiopod assemblages can be identified on most of the main palaeocontinents prior to the Ordovician radiation of the Palaeozoic Evolutionary Fauna.