Abstract A Cambro-Ordovician palaeogeographical restoration of the southwestern European margin of Gondwana is proposed based on the relative positions of Variscan tectonostratigraphic units. Four palaeogeographical proximal–distal transects are recognized and comprise: (i) the Cantabrian, West Asturian-Leonese, Central Iberian/Central Armorican and Ossa-Morena/North Armorican zones and domains of the Iberian and Armorican massifs, respectively; (ii) the South Armorican Domain and its lateral prolongation into the Thiviers-Payzac unit and the Occitan Domain, including the transect from the Axial, southern and northern Montagne Noire, and the Albigeois-southern Cévennes unit; (iii) the southern and northern sides of the Canigó Massif in the Eastern Pyrenees; and (iv) the External Zone and the External and Internal nappes of Sardinia. Two geodynamic scenarios are recognized controlled by the presence/absence of: (i) the Furongian–Early Ordovician (Toledanian or ‘lacaune normande’) break-up unconformity across the Ossa-Morena/North Armorican and Central Iberian/Central Armorican belts; (ii) the Early–Late Ordovician (Sardic) Phase across the Occitan and Pyrenean domains and SW Sardinia; and (iii) the migration of peaks in trilobite and cinctan (echinoderm) diversity. Other similar palaeogeographical shifts are recognized in zircon provenance patterns, the occurrence of climatically sensitive subtropical facies and mineral indicators across platform–basinal transects along the Gondwana margin. This multidisciplinary framework is proposed as a preliminary step in the quest to produce more tightly constrained Early Paleozoic reconstructions along southwestern Europe.

Abstract The distribution of all known Cambrian echinoderm taxa, encompassing both articulated specimens and taxonomically diagnostic isolated ossicles, is documented for the first time. The database described by 2011 comprises 188 species recorded from 65 formations from around the world. Formations that have yielded articulated echinoderms are unequally distributed in space and time. Only Laurentia and West Gondwana provide reasonably complete records at the resolution of Stage. The review of the biogeographical distributions of the eight major echinoderm clades shows that faunas from Laurentia and Northeast Gondwana (China and Korea) are distinct from those of West Gondwana and Southeast Gondwana (Australia); other regions are too poorly sampled to make firm palaeobiogeographical statements. Analysis of alpha diversity (species per formation) shows that diversity rose initially to Cambrian Stage 5, declined into Guzhangian and Paibian before returning to Stage 5 levels by the end of the Cambrian. This pattern is replicated in Laurentia and West Gondwana. We show that taxonomically diagnostic ossicles found in isolation typically occur significantly earlier than the first articulated specimens of the same taxa and provide important information on the first occurrence and palaeobiogeographical distribution of key taxa, and of the phylum as a whole. Supplementary material: Articulated Cambrian echinoderms and Isolated plates of Cambrian echinoderms are provided at: http://www.geolsoc.org.uk/SUP18668

Abstract Palaeobiogeographical data on Cambrian trilobites obtained during the twentieth century are combined in this paper to evaluate palaeoceanographic links through c. 30 myr, once these arthropods biomineralized. Worldwide major tectonostratigraphic units are characterized at series intervals of Cambrian time and datasets of trilobite genera (629 for Cambrian Series 2, 965 for Cambrian Series 3, and 866 for the Furongian Series) are analysed using parsimony analysis of endemicity. Special attention is given to the biogeographical observations made in microcontinents and exotic terranes. The same is done for platform-basinal transects of well-known continental margins. The parsimony analysis of endemicity analysis resulted in distinct palaeogeographical area groupings among the tectonostratigraphic units. With these groupings, several palaeobiogeographical units are distinguished, which do not necessarily fit the previously proposed biogeographical realms and provinces. Their development and spatial distributions are broadly controlled by Cambrian palaeoclimates, palaeogeographical conditions (e.g. carbonate productivity and anoxic conditions) and ocean current circulation. Supplementary material: Global dataset of Cambrian Epoch 2 (A), Cambrian Epoch 3 (B) and the Furongian Epoch (C) trilobite genera are provided at: http://www.geolsoc.org.uk/SUP18669

Abstract Carbonate productivity and glaciomarine deposits of the Ordovician–Silurian transition display different sedimentary architectures in the Iberian and Hesperian Chains of NE Spain, as a result of quiescent and active extensional tectonics on platforms fringing North Gondwana. The late Katian carbonate productivity of the Iberian platform reflects the onset of bryozoan–pelmatozoan meadows and mud-mound complexes throughout an intra-shelf ramp, whereas carbonate nucleation of prominent carbonate factories took place on the top of isolated palaeo-highs in the Hesperian platform. In both cases, the end of carbonate productivity is associated with glacioeustatic regression, subaerial exposure and karstification, pre-dating widespread precipitation of iron ore deposits in the vicinity of palaeo-highs. The Hirnantian glacioeustatic transgression is represented lithostratigraphically by the Orea Formation. In the Iberian platform, the formation consists of two distinct depositional sequences bounded by the progradation of conglomeratic channels, and is dominated by the record of massive and crudely stratified diamictites, with tabular geometries and deposited subaqueously as ‘rain-out’ facies. In contrast, the Hesperian platform is rich in disrupted diamictites, which form strongly deformed units interpreted as submarine slumps associated with active synsedimentary faults. In both cases, the anomalous occurrence of massive diamictites, rich in boulder- to sand-sized carbonate dropstones, and displaying rapid variations in density and size, suggests that distinct iceberg ‘drift lanes’ existed, indicating current activity in the open sea.

Abstract An interval of episodic carbonate productivity, lithostratigraphically recognized as the ‘Calcaires inférieurs’ (upper member of the Adoudou Formation), took place across the Neoproterozoic–Cambrian transition onlapping the western Saghro inlier, Morocco. Sedimentation of the ‘Calcaires inférieurs’ was highly variable: in relatively stable substrates, a peritidal-dominated mixed platform is recorded where deposition was primarily controlled by autocyclic processes and accommodation space availability, whereas, in unstable substrates, the tectonic activity associated with the inherited block-faulting basement led to deposition of complex slide sheets composed of penecontemporaneous isoclinal folds and disrupted strata. The uppermost part of the ‘Calcaires inférieurs’ displays a negative δ 13 C shift reaching values of −6.5‰. This shift may represent the δ 13 C excursion to −6‰ that marks the Neoproterozoic–Cambrian boundary in the western Anti-Atlas. Two volcanic episodes bracketed the carbonate productivity. They consist of lower basaltic flows and an upper rhyolitic ignimbrite, with a SiO 2 gap between 52 and 74 wt%. The basic rocks resemble those of tholeiitic magmas in continental rifts. The felsic rocks show high light to heavy rare earth element abundances and negative Nb, Ta, P and Ti anomalies, and were probably generated as a result of either fractional crystallization coupled with relative crustal contamination, or from a different magmatic source. The lower basic flows of tholeiitic affinity predated and geochemically differ from the alkaline magmatism of the Alougoum volcanic complex (Boho jbel) that surrounds the neighbouring Bou-Azzer inlier.

Abstract One unresolved conceptual problem in some Palaeozoic sedimentary strata is the boundary between the concepts of “shell concentration” and “reef”. In fact, numerous bioclastic strata are transitional coquina–reef deposits, because either distinct frame-building skeletons are not commonly preserved in growth position, or skeletal remains are episodically encrusted by “stabilizer” (reef-like) organisms, such as calcareous and problematic algae, encrusting microbes, bryozoans, foraminifers and sponges. The term “parabiostrome”, coined by Kershaw, can be used to describe some stratiform bioclastic deposits formed through the growth and destruction, by fair-weather wave and storm wave action, of meadows and carpets bearing frame-building (archaeocyaths, bryozoans, corals, stromatoporoids, etc.) and/or epibenthic, non-frame-building (e.g. pelmatozoan echinoderms, spiculate sponges and many brachiopods) organisms. This paper documents six Palaeozoic examples of stabilized coquinas leading to (pseudo)reef frameworks. Some of them formed by storm processes (generating reef soles, aborted reefs or being part of mounds) on ramps and shelves and were consolidated by either encrusting organisms or early diagenesic processes, whereas others, bioclastic-dominated shoals in barrier shelves, were episodically stabilized by encrusting organisms, indicating distinct episodes in which shoals ceased their lateral migration.

Abstract Shelled phosphorites of Early Cambrian age are common in the Avène-Mendic autochthonous unit (Marcory Formation) and the Mélagues nappe ( “Heraultia beds” of the Lastours Formation), northern Montagne Noire (France). Palaeogeographically, the concentration of phosphate took place along the shelf edge between a stable inner platform (southern Montagne Noire) and an unstable slope-to-basin sea floor preserved in the northern Montagne Noire. Petrography, back-scattered SEM (scanning election microscopy) and elemental mapping by EDS (energy dispersive system) show that the phosphorites were generated by repeated alternations of low sedimentation rates and condensation forming hardgrounds, in situ early diagenetic phosphogenesis, winnowing and polyphase reworking of previously phosphatized skeletons and hardground-derived clasts. The succession of repeated cycles of sedimentation, phosphate concentration and reworking led to multi event phosphate deposits rich in allochthonous particles. Associated accumulations of exhumed and reworked pyrite clasts reflect final deposition in a mainly dysaerobic substrate.

Abstract All bioherms from the Early Cambrian (Botoman) Matoppa Formation of the Nebida Group in SW Sardinia were previously thought to be dominated by Epiphyton. However, at “La Sentinella” (Serra Scoris Hill), they are composed of Girvanella, Razumovskia, Botomaella and Renalcis, with Epiphyton and archaeocyaths as accessory components. This association forms two unusual types of crust boundstone, consisting of stacked flat or curved crusts and saucer-like archaeocyaths delimiting shelter cavities. Dendrolitic Renalcis archaeocyath–cement boundstone caps the bioherm. Analysis of the La Sentinella reef complex and comparison with similar constructions from Mongolia (Zuune Arts, Salaany Gol), Nevada (Stewart's Mill, Battle Mountain), Mexico (Sonora) and China (Tianheban Formation) suggest that episodic deposition of fine-grained siliciclastic or carbonate sediment followed by periods of non-deposition enabled the calcimicrobial rafts and crusts to colonize the substrate and then provide synoptic relief for the development of a dendrolitic Renalcis–cement framework. “La Sentinella” is one of the rare examples of Cambrian reef complex displaying community replacement, from an initial stage of thrombolitic and/or flat-stacked microbial crusts on a muddy substrate to an arched microbial crust system, to a more resistant Renalcis–cement boundstone. Such bioherms reflect an open-shelf, shallow-marine environment of increasing energy.

Abstract Exposures of the Botoman (Lower Cambrian), Lemdad and Issafen formations on the Lemdad syncline, southern High Atlas, provide an excellent example of the interactions between tectonic events, magmatic activity and carbonate productivity. The major factors that controlled the nucleation of carbonate factories on the Botoman High Atlas platform were: (i) synsedimentary tectonism, as normal faulting resulted in tilting of fault blocks causing irregular topographies and subsequent sharp erosion; (ii) volcanism, because pyroclastic influx smothered carbonate factories except in distal areas of the platform or during quiescent episodes of volcanic activity; and (iii) the influence of successive shoaling parasequences. The Botoman reefs exhibit a wide range of external morphologies, including tabular (biostromes) and domal (bioherms and patches) boundstones, which do not exceed 3.5 m of thickness. Although archaeocyathan–microbial reefs only developed under clear-water conditions, microbial reefs grew also under turbid-water conditions. Domal and digitate stromatoids, Girvanella crusts, Epiphyton bushes and thromboid–stromatoid intergrowths document the ability of some microbial communities to develop heterotrophic strategies when submitted to a moderate terrigenous input. Turbidity was a major ecological factor that constrained development of filter/suspension-feeder and phototrophic organisms, but not necessarily of benthic non-phototrophic microbial communities.

Abstract In the abandoned slate quarry of Guernanic, Gourin (Morbihan, France), a single horizon (Upper Member of the Schistes de Postolonnec Formation) has yielded an exquisitely preserved Llandeilian (Middle Ordovician) echinoderm assemblage composed of the ophiuroid Taeniaster armoricanus sp. nov. and the mitrate Mitrocystella incipiens. These two groups of echinoderms represent the first fossils formally described from the Middle Ordovician of the Gourin area. The brittlestar T. armoricanus sp. nov. is the third and oldest ophiuroid reported so far in the Palaeozoic of the Armorican Massif. The mitrate Mitrocystella is also described for the first time from western Brittany. Taphonomic features of this ophiuroid–stylophoran aggregation suggest that it probably corresponds to the rapid burial of a life assemblage in an otherwise quiet and moderately deep setting (shelf) below, but close to, storm wave base. This echinoderm association represents the oldest evidence for a gregarious mode of life for ophiuroids, as well as the oldest indisputable example of a mixed ophiuroid–stylophoran meadow.

Abstract Silurian reefs are well known to comprise frame-building corals, stromatoporoids and algae, but also a range of calcimicrobial components and micritic sediments of possible microbial origin. The margins of Wenlock patch reefs tend to have diffuse edges that grade into the surrounding bedded facies because of talus shed from the reefs. However, portions of patch reefs show sharp-bounded reef margins, with bedded limestones terminating abruptly against the reef edge. Examples of sharp boundaries where the reef comprises only carbonate mudstone–wackestone with poorly-defined gross fabric, and containing no metazoan framework, have been found in Wenlock patch reefs the UK and Gotland. Although in some cases the micrite may demonstrate a peloidal structure, in others there is no clear structure, broadly fitting the aphanitic (structureless) type of fabric found in leiolites (suspected microbial facies that show no structure). The fabrics are interpreted to have been formed by microbial mediation of micrite precipitation as part of reef construction, and are therefore automicrites. In all cases the sharp reef edges indicate coherence of the micritic fabric, interpreted as a lithified wall against which bedded limestones were deposited. This arrangement supports published interpretations of pronounced topography of Wenlock patch reefs, and shows the presence of steep, vertical and, possibly, overhanging reef margins, formed prior to bedded sediment accumulation. Thus, there is likely to have been a time interval between reef formation and deposition of bedded sediments, possibly caused by reef upward growth in transgressive settings, followed by catch-up of bedded limestone deposition.

Abstract Palaeozoic sediments of Austria are separated by the Periadriatic Fault into Eastern Alpine (Upper, Middle and Lower Austroalpine) and Southern Alpine units. We herein present six case studies showing up the different development of shallow-marine communities with special regard to carbonate factories and shell pavements occurring in both regions during the Siluro-Devonian time span. Upper Silurian-Upper Devonian deposits of the Eastern Alps comprise accumulations of serpulid tubes (Southern Burgenland) and Septatrypa pavements, Amphipora mounds, coral-stromatoporoid–biostromes and Stachyodes–auloporoid beds regarded as pioneer reef communities (Graz Palaeozoic), respectively. Lower Silurian strata of the Southern Alps consist of pelagic sediments persisting to the Upper Silurian and therefore differ from contemporaneous successions in the Eastern Alps. Intercalated in Ludlow orthocerid limestone beds Cardiola pavements appear (Carnic Alps). Within the Lower Devonian sequence, mounds were built by baffling calcareous algae and tabulozoan communities. Coral–stromatoporoid patch reefs occur during the Pragian, Givetian and Frasnian stages.