Abstract The Ordovician was a key period in the biological and geological history of the Earth. ‘A Global Synthesis of the Ordovician System’ is presented in two volumes of The Geological Society, Special Publications series. The first volume (SP532) covers general aspects of the Ordovician and also includes the syntheses of the Ordovician successions of Europe. To provide a comprehensive global overview, this second volume (SP533) represents a journey through the Ordovician System around the world. Reviews of the Ordovician of North America include syntheses of Alaska, Greenland, Canada, the USA and Mexico, whereas the South American Ordovician is summarized in a specific chapter related to Argentina and neighbouring countries. The Ordovician System of Africa is presented in chapters covering the north and the south of the continent where significant Ordovician successions occur. Australia and New Zealand, as well as Antarctica, are visited in separate chapters. Asia provides the most complex Ordovician successions that are reviewed in chapters covering Turkey and the Levant region, the Middle East, Central Asia, Kazakhstan, India, SE Asia, China, Korea, and Japan. Our journey covers a great number of locations but, with many successions still to be fully described, our knowledge of the Ordovician of the world remains incomplete.

Abstract Evidence of Early Ordovician deposition and intrusion in East Antarctica is best known from the Ross Orogen, postdating the 495–489 Ma Ross Orogeny. Here, c. 490–475 Ma granites (with related dykes and sills) of the Granite Harbour Intrusives represent roots of a continental margin arc. Detrital zircon grains in the upper Byrd Group (Central Transantarctic Mountains) are of comparable Early Ordovician age. Contemporaneous fossils are rare. In northern Victoria Land they include latest Cambrian to earliest Ordovician conodonts and microbrachiopods in allochthonous limestones of the Handler Formation (Robertson Bay Group) in the Robertson Bay Terrane, and probable Early Ordovician trace fossils in the Camp Ridge Quartzite of the Leap Year Group in the Bowers Terrane. In the Shackleton Range of Coats Land, West Antarctica, the Blaiklock Glacier Group contains a diverse ichnofossil fauna of probable Ordovician age associated with undescribed bivalved arthropods and segmented crustacea. The Swanson Formation of the Ross Province in Marie Byrd Land (correlated with the Robertson Bay Group of the Ross Orogen) is a turbiditic unit dominated by quartz-rich sandstones. Its Ordovician age is based on a post-depositional whole rock K–Ar metamorphic age of 448–444 Ma, with detrital zircon grains indicating a late Cambrian maximum depositional age.

Abstract The stratigraphic overview presented in this chapter substantially updates and revises the last major review of the Ordovician rocks of Australia and New Zealand published 40 years ago. In the western two-thirds of the present-day continent of Australia, Ordovician sedimentary rocks are restricted to intracratonic basins. The Canning Basin (Western Australia) and Amadeus Basin (central Australia) contain the best known Lower and Middle Ordovician shallow marine successions. The eastern third of the continent, known as the Tasmanides, comprises multiple orogens (i.e. Delamerian, Lachlan, New England, Thomson, Mossman) that formed along the convergent East Gondwana Margin. As a result, volcanic and intrusive rocks are much more common in these orogens than in the intracratonic basins. Their deep-water depositional environments span 31 graptolite biozones. Slope and basinal siliceous sedimentary rocks are constrained by a newly defined set of 12 conodont biozones, complementing the conodont biostratigraphic scheme refined for shallow-water environments from the basal boundary of the Ordovician to the latest Katian. In some places, these conodont biozones are integrated with radiometric ages from tuff interbeds (e.g. Canning Basin). Ordovician graptolitic strata in the Buller Terrane of New Zealand share palaeogeographic links with those in the Bendigo Zone of the western Lachlan Orogen.

Abstract The Ordovician was a key period in the biological and geological history of the planet. ‘A Global Synthesis of the Ordovician System’ is presented in two volumes of The Geological Society, Special Publications . This first volume (SP532) charts the history of the Ordovician System and explores significant advances in our understanding of the period's biostratigraphy, including more precise calibration of its timescale with tephra chronology and regional alignments using astrochronology and cyclostratigraphy. Changes in the world's oceans, their shifting currents and sea levels, the biogeography of their biotas and the ambient climate are described and discussed against a background of changing palaeogeography. This first volume also includes syntheses of Ordovician geology of most European countries, including historical key areas, such as the British Isles, Baltoscandia and Bohemia.

Abstract The biogeographical patterns shown by Ordovician linguliform and craniiform brachiopods are greatly influenced by their dominance in low-diversity associations in marginal environments. This is particularly evident in the Early Ordovician, when linguliform-dominated dysaerobic assemblages are widely distributed along the deep shelves of Gondwana, the Kazakhstanian terranes and in Baltica. By the Darriwilian, micromorphic linguliforms are characteristic components of the pantropical climatic-controlled faunas of Laurentia, Cuyania and Kazakhstanian terranes, which – in spite of separation by extensive oceans – retain a distinct similarity. Analysis of craniiform biogeographical distribution is impeded significantly by the poor state of craniide taxonomy and lack of reliable data from most regions. However, in general their biogeographical dispersion is similar to other groups of the Palaeozoic Evolutionary Fauna. Unlike the linguliforms, which are important members of the Cambrian Evolutionary Fauna, there is no convincing Cambrian craniiform record; they may have evolved and dispersed from Gondwana and associated microcontinents and island arcs. The earliest well-established record is from the late Tremadocian of temperate to high-latitude peri-Gondwana. During most of the Ordovician, they have a peri-Iapetus distribution. They are very rare or absent in tropical Gondwana, South China and Kazakhstanian terranes and are not yet documented from Siberia. The trimerellides probably evolved in tropical peri-Gondwanan island arc settings. Their dispersion and major features of biogeography mirror those of atrypides. Gold Open Access: This article is published under the terms of the http://creativecommons.org/licenses/by/3.0/ .

Abstract The phylogeographical evolution and the consequent changing distribution and diversity of rhynchonelliform brachiopods through the Ordovician are linked to the dynamic palaeogeography of the period. The Early Ordovician (Tremadocian and Floian) is characterized by globally low-diversity faunas with local biodiversity epicentres, notably on the South China Palaeoplate; low-latitude porambonitoid-dominated faunas with early plectambonitoid and clitambonitoid representatives, as well as high-latitude assemblages mostly dominated by orthoids, can be recognized, but many taxa are rooted in Late Cambrian stocks. The Early Ordovician displays a steady increase in rhynchonelliformean biodiversity, which was mostly driven by the increasing success of the Porambonitoidea and Orthoidea, but the billingsellids and early plectambonitoids also contributed to this expansion. During the Early to Mid Ordovician (Dapingian–Darriwilian), marine life experienced an unprecedented hike in diversity at the species, genus and family levels that firmly installed the suspension-feeding benthos as the main component of the Palaeozoic fauna. However, this may have occurred in response to an early Darriwilian annihilation of existing clades, some of which had been most successful during the Early Ordovician. New clades rapidly expanded. The continents were widely dispersed together with a large number of microcontinents and volcanic arcs related to intense magmatic and tectonic activity. Climates were warm and sea-levels were high. Pivotal to the entire diversification is the role of gamma (inter-provincial) diversity and by implication the spread of the continents and frequency of island arcs and microcontinents. The phylogeographical analysis demonstrates that this new palaeogeographical configuration was particularly well explored and utilized by the strophomenides, especially the Plectambonitoidea, which radiated rapidly during this interval. The porambonitoids, on the other hand, were still in recovery following the early Darriwilian extinctions. Orthides remained dominant, particularly at high latitudes. Biodiversity epicentres were located on most of the larger palaeoplates, as well as within the Iapetus Ocean. Provincial patterns were disrupted during the Sandbian and early Katian with the migration of many elements of the benthos into deeper-water regimes, enjoying a more cosmopolitan distribution. Later Katian faunas exhibit a partition between carbonate and clastic environments. During the latest Katian, biogeographical patterns were disrupted by polewards migrations of warm-water taxa in response to the changing climate; possibly as a consequence of low-latitude cradles being developed in, for instance, carbonate reef settings. Many clades were well established with especially the strophomenides beginning to outnumber the previously successful orthides, although this process had already begun, regionally, in the mid to late Darriwilian. At the same time, atrypoid and pentameroid clades also began to radiate in low-latitude faunas, anticipating their dominance in Silurian faunas. The Hirnantian was marked by severe extinctions particularly across orthide-strophomenide clades within the context of few, but well-defined, climatically controlled provincial belts. Supplementary material: The individual localities and a reference list for the data sources are provided at: http://www.geolsoc.org.uk/SUP18667

Abstract The biogeographical distribution of Ordovician and Silurian gastropods, monoplacophorans and mimospirids has been analysed on a generic level. The dataset contains 334 genera and 2769 species, yielding 1231 records of genera with 2274 occurrences worldwide. There is a bias towards eastern Laurentia, Baltica and Perunica records. Some 53.1% of the records are Ordovician. The study demonstrates that these molluscs are well suited to being used to improve understanding of Ordovician and Silurian biogeographical provinciality. Specific points are that: a Lower Ordovician assemblage is evident in Laurentia; the fauna of the Argentinean Precordillera is Laurentian until the Darriwilian, when taxa are shared with North China; Late Silurian gastropods from the Alexander terrane (SE Alaska) are unknown in Laurentia, but support a rift origin of this terrane from NE Siberia; Perunica, Ibero-Armorica and Morocco cluster together throughout the Ordovician but Perunica and Morocco are closer; Darriwilian–Sandbian deep-water Bohemian taxa occur in Baltica; a Laurentian–Baltica proximity is unsupported until the Silurian; Siberia clusters with North China and eastern Laurentia during the Tremadocian–Darriwilian; during the Gorstian–Pridoli Siberia clusters with the Farewell and Alexander terranes; North China may have been close to Laurentia and the Argentinean margin of Gondwana; and the affinity of Tarim taxa is problematic.