Abstract More than 30 well-preserved pterosaurian trackways have been excavated from the site of Crayssac (Lower Tithonian, southwestern France), along with hundreds of isolated imprints. The general pattern of pterosaurian trackways is described and corresponds to the morphology of the ichnogenus Pteraichnus Stokes 1957 . A crocodilian origin for Pteraichnus is rejected. The morphology of the pes and manus prints is described and discussed, and exhibits a great dynamic variability. The three manus digit prints are confirmed as the marks of digits I—III, but the high angle of divarication may reflect strong digit abduction as well as a backward folding of the digit III. The manus gait-width is as much as 3 × the pes gait-width. The narrowest manus trackways corresponds to low-velocity walking in a semi-erect stance, whereas the widest manus trackways corresponds to higher velocity walking (perhaps even running), with a subhorizontal body.