Abstract Owing to the increasing availability of data for many fossil groups and a generally accepted palaeogeographical configuration, palaeontologists have been able to develop progressively more robust palaeobiogeographical scenarios for the spatial distributions of Ordovician marine faunas. However, most research in Early Paleozoic palaeobiogeography centres on data derived from extensively studied localities in North America and Europe. Thus, clear patterns are emerging of regional biogeography for these areas. However, the fragmentary nature of data from other regions hinders the development of a detailed understanding of palaeogeographical schemes of many clades at the global level. Provincial patterns are now available for several fossil groups, but the global coverage remains generally fragmentary. Palaeobiogeographical investigations were traditionally focused on better understanding of palaeogeographical scenarios and often employed quantitative analyses of faunal similarity. More recently palaeobiogeographical analyses have expanded to investigate questions such as the location and pace of speciation and macroevolution together with macroecological change. For example, studies on the evolution of speciation levels in the frame of the taxonomic radiation of the Great Ordovician Biodiversification are now available. Future investigations, including modelling, will provide more integrative, global patterns of provincialism, including the location of Ordovician biodiversity hotspots and the recognition of latitudinal diversity gradients.

Abstract More than 30 species of tergomyan, gastropod, bivalve and cephalopod molluscs are described from the Late Ordovician of central and eastern Anti-Atlas, Morocco. For the cephalopods this represents the first systematically known taxa of the region. Tergomyans and gastropods are most common in the more shaly part of the Lower Ktaoua Formation, while bivalves are more frequent in the sandy part of the Lower Second Bani Formation. A southern Gondwana aspect is evident for the tergomyan, gastropod and bivalve assemblages, sharing many taxa with Bohemia, while no clear signal is present for the cephalopods. The latter reflects insufficient knowledge of the fauna of large parts of these areas. The widely recognized Holopea ? antiquata is transferred to the genus Radvanospira gen. nov.; other new gastropod taxa include Allossospira gen. nov., Tritonophon grandis sp. nov., Radvanospira baniensis sp. nov. and Lophospira latilabra sp. nov. Most of the diverse bivalve fauna and the few cephalopods are left in open nomenclature but new taxa include the bivalve Praenucula pojetai sp. nov. and the cephalopods Wadema tattai sp. nov. and Tafadnatoceras tiouririnense gen. et. sp. nov. The Late Ordovician bivalves from Morocco are dominated by pteriomorphs and protobranchs, inhabiting infaunal, semi-infaunal and epifaunal niches.

Abstract Cephalopods have their earliest occurrence in Late Cambrian shallow-water carbonates on the North China Platform and rapidly dispersed across the globe within the latest Cambrian. Latest Cambrian and initial Ordovician cephalopod occurrences are restricted to the palaeotropical realm. The Ordovician records a unique morphological diversification and expansion of cephalopod habits and habitats which is expressed in a massive morphological diversification and unique palaeogeographical patterns of dispersal. The Ordovician cephalopod diversification was a complex process of appearance and disappearance of higher groups with a specific palaeogeographical signature and a clear selective component. A general Ordovician trend showed decreasing evolutionary turnover rates, increasing number of widespread genera, decreasing proportion of endemic genera, and decreasing beta-diversity. This is interpreted as a result of an increasing ecosystem stability during this time interval.