A bstract Upper Cenomanian limestones exposed in southern and western Istria, Croatia, are rich in radiolitid remains and also containdinosaur footprints. As documented in measured sections at Lovrecica, Ladin Gaj, Grakalovac, and Fenoliga, as well as in a new sectionat Karigador, rudists and dinosaur tracks occur together in peritidal sediments that are organized into shallowing-up cycles. These cycleswere deposited in the interior of the carbonate platform and are commonly capped by intertidal laminites. At these localities, the rudistsare elevator types and mud-supported sediment dwellers that thrived in the soft substrate where dinosaurs made footprints. Dinosaurtracks are preserved in both types of subtidal limestones, and radiolitid congregations are laterally discontinuous in foraminiferal-peloidal limestone. The presence of dinosaur footprints on the upper bedding plane of rudist floatstone implies that the Istrian part of the Late Cenomanian carbonate platform was a shallow, protected platform interior. Dinosaurs strode across the muddy bottom of the shallow-water platformin water depths equivalent to their estimated hip heights. These dinosaur-track-rich levels in Upper Cenomanian shallow marinelimestones of the Adriatic carbonate platform underlines the relevance of dinosaur footprints as a tool for paleoenvironmental reconstruction.