The Woolhope inlier, near Ledbury, Wales, has several zones of fragmentary beds in the Ludlovian (Silurian) strata that have yielded thelodont and acanthodian denticles. Thelodont remains have also been found in the Usk and Lye inliers at the base of the Dayia shales and very probably occur at this horizon over the entire area. While concentrations of fish remains are not found in the Llandovery or Wenlock series, they are undoubtedly present though scattered.

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