In north Staffordshire, central England, the base of the Tenuis zone (Carboniferous) has formerly been placed at the base of the Keele beds and above the Newcastle-under-Lyme beds. It has been previously suggested that part of the Newcastle beds may belong to the Tenuis zone. The discovery of Anthraconauta tenuis in strata just above the basal limestone of the Newcastle beds confirms the previous assumption.

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