Two hundred and eighty specimens of paradoxidids from two localities in the Holy Cross Mountains (Poland) have been reanalysed using morphometric techniques. Revision of the dataset provided evidence for the presence of two endemic taxa: Acadoparadoxides kozlowskii (Orłowski, 1959) and Acadoparadoxides samsonowiczi (Orłowski, 1959), both belonging to the earliest group of Acadoparadoxides, initially considered to be present only in Gondwanan successions. Thus, this is the first description of the members of this group outside West Gondwana. The taxonomic revision, and the presence of representatives of the earliest acadoparadoxidines coupled with the absence of taxa typical of Scandinavia that were previously described from this locality have resulted in the modification of the biostratigraphic scheme hitherto used in the Holy Cross Mountains. The newly established A. samsonowiczi – A. kozlowskii Assemblage Zone is correlated with the Morocconus notabilis Zone of Morocco, and thus represents the Cambrian Series 2 and 3 boundary interval. Links with West Gondwana challenge the existing palaeogeographic interpretations for the southern part of the Holy Cross Mountains and point to an urgent need to revise the position of the Małopolska Block during the middle Cambrian.