We describe Eocene fossils of the tillodont Trogosus from the Allenby Formation in Princeton, British Columbia (B.C.), as well as teeth of Brontotheriina from the lower Australian Creek Formation near Quesnel, B.C. These fossils represent the only occurrence of Tillodontia and Brontotheriidae in B.C. Further, the presence of the largest species of TrogosusT. latidens — as well as a smaller species identified only as Trogosus sp. supports a late early – early middle Eocene (Bridgerian) age for the Vermilion Bluffs Shale of the Allenby Formation. Based on their morphology and large size, the teeth referred here to Brontotheriina represent one of the larger, more derived brontothere genera, and suggest a Uintan–Chadronian (middle–late Eocene) age range for the lower Australian Creek Formation that is consistent with radiometric ages of underlying volcanic rocks. Paleobotanical data from sediments correlative to those that produced these Eocene mammal fossils suggest they inhabited forested landscapes interspersed with swamps and open water environments, under mild and wet temperate climates (mean annual temperature (MAT) ∼10–16 °C; cold month mean temperature (CMMT) −4–4 °C; mean annual precipitation (MAP) >100 cm/year). These mixed conifer–broadleaf forests included tree genera typical of modern eastern North American forests (e.g., Tsuga, Acer, Fagus, and Sassafras), together with genera today restricted to east Asia (e.g., Metasequoia, Cercidiphyllum, Dipteronia, and Pterocarya). The paleobotanical evidence is consistent with the hypothesized habitats of both tillodonts and brontotheres.

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