Two samples of late Paleozoic grit and Late Mississippian quartzite–chert conglomerate collected from southeastern Yukon–Tanana terrane (YTT) — a composite thrust sheet resting structurally above North American parautochthonous strata and intervening imbricate sheets of the late Paleozoic oceanic Slide Mountain terrane — yielded, respectively, 89 and 74 concordant or nearly concordant (<20% discordant) U–Pb ages on single detrital zircons. They provide constraints on the provenance of this allochthonous pericratonic terrane. Zircons in the grit range from 1770 to 2854 Ma, with a well-defined Early Proterozoic peak between 1800 and 2100 Ma. Precambrian zircons in the conglomerate also show a dominant peak between 1800 and 2100 Ma and smaller peaks between 2200 and 3200 Ma, with a few older grains, and younger grains with ages of 998, 1219, 1255, 1256, and 1417 Ma. The conglomerate also yielded three Devonian grains, with ages of 366 ± 23, 373 ± 12, and 379 ± 23 Ma. Their ages are approximately coeval with the oldest felsic to intermediate arc- and rift-related magmatism in the YTT. The age spectra from southeastern YTT units compare closely with those from Mississippian and older pericratonic units in the Coast Mountains, confirming correlations previously made on lithologic grounds. They also strongly resemble detrital zircon populations from craton-derived Paleozoic units of the northern North American autochthon. This robust U–Pb data set lends support to the idea that YTT once formed part of the outer, active margin of the North American continent, prior to Mississippian rifting and marginal ocean basin development.