Abstract

The hitherto unknown female fore wing (tegmen) of the Paleocene orthopteroid, Albertoilus cervirufi Kevan and Wighton (Grylloptera; Hagloidea), has been discovered at the type locality of the species. This has facilitated a more critical assessment of the systematic affinities of Albertoilus. The relationships of the genus appear to lie, on the one hand, with the central Asiatic Late Jurassic prophalangopsine genus Aboilus (as had previously been indicated) and, on the other, with the western North American Oligocene genus Palaeorehnia. The type species of the latter, P. maculata Cockerell, 1908, is shown to be a junior synonym of "Cymatomera" maculata Scudder, 1890, described earlier from the same site. This has led to the new nominal combination Palaeorehnia maculata (Scudder) and a more definite taxonomic assignment of that species. The family group name Palaeorehniinae falls as a junior synonym of Prophalangopsinae. A new subfamily name Zeuneropterinae (Stenopelmatoidea, Gryllacrididae) is proposed for the genus Zeuneroptera Sharov. It is concluded that recent North American Prophalangopsidae (Cyrtophyllitinae) are derived directly from eastern Asiatic forebears that crossed the Beringian land bridge, and that the earlier North American members of the family (Prophalangopsinae) became extinct in the Tertiary. The family Phasmomimidae was recently indicated by us as having been previously unrecorded for North America, or later than the Early Cretaceous. It had in fact been reported erroneously as a member of the order Plecoptera from lower Upper Cretaceous strata of Labrador in 1969 (undescribed material of the same approximate age is also known from the eastern U.S.S.R.). The species in question, Palaeopteron complexum Rice, is very similar to Phasmomimella paskapoensis Kevan and Wighton, and it is possible that their respective genera might be synonymized, despite a long time interval separating their known occurrences.The tettigoniid subfamily name Pseudotettigoniinae should be attributed to Sharov, not to Kevan and Wighton. It is conceivable that the unnamed Paleocene tettigoniid described by the latter authors may be in some way related to the Oligocene pseudottigoniine genus Lithymnetes Scudder.

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