Brachiopod and trilobite faunal distributions indicate that the Iapetus Ocean was still wide enough to inhibit migration in the Middle and Late Ordovician. The presence of Silurian and Lower Devonian calc-alkaline rocks suggests that ocean crust was still being subducted long after the end of the Ordovician and that the Iapetus Ocean did not finally close in Newfoundland until the Acadian Orogeny. The Reach Fault divides successions containing different Lower Palaeozoic faunas; to the west, typical North American faunas occur in New World Island (Cobb's Arm Limestone), while to the east the rocks of the Gander region appear to have been attached to the Avalon Peninsula, with its European Lower Palaeozoic faunas, since the Early Ordovician. It is concluded that the Reach Fault marks the suture where the Iapetus Ocean closed at the end of the Early Devonian. This line probably extends across Newfoundland to the south of Buchans, and links up with the Cape Ray Fault in the southwest of the island. An Ordovician fauna from the Davidsville Group of the Gander area is illustrated for the first time; it is not clearly definitive of any faunal province.