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Abstract

Three species of artiodactyls (Diacodexis sp. cf. D. secans, Simpsonodus chacensis [new genus and species], and Wasatchia grangeri) are reported from the San Jose Formation, New Mexico; these three and an additional four taxa (Simpsonodus sp., W. pattersoni, Bunophorus sinclairi, and Hexacodus pelodes) occur in the Debeque Formation, Colorado. The new diacodexeid genus, Simpsonodus, includes D. chacensis and W. lysitensis. Wasatchia pattersoni, new species, the most derived known species of Wasatchia, is morphologically suitable as an ancestor to Bunophorus, with which it overlaps stratigraphically. Diacodexis sp. cf. D. secans is morphologically and apparently phylogenetically intermediate between D. secans, on the one hand, and Hexacodus and Antiacodon on the other. The early Eocene record of diacodexeids is robust enough to indicate: (1) the gradual and continuous divergence of D. secans and Diacodexis sp. cf. D. secans in the mosaic distribution of derived character states; and (2) from these taxa, respectively, the gradational origin of other diacodexeids/leptochoerids and antiacodontids/homacodontids. In these instances, microevolutionary processes appear to account for the macroevolutionary pattern.

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