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The Death Valley system (southeastern California and southwestern Nevada) contains a locally endemic aquatic biota that has long been the subject of compelling biogeographic speculation, yet it remains little studied phylogenetically. Springsnails (Hydrobiidae: Pyrgulopsis) are one of the most diverse elements of this fauna, and they are thought to have evolved in association with late Tertiary rearrangements of landscape and drainage. We assembled a molecular phylogeny for this fauna to investigate its evolutionary development in relation to regional geological history. Sequences for two mitochondrial genes were obtained from 80 populations representing 13 of the 14 Death Valley system springsnail...

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