The early Late Devonian (early Frasnian) Alamo Impact targeted an oceanic, off-platform site in southern Nevada, excavating a crater with a final diameter of 44–65 km. The original crater is now dismembered and buried beneath younger rocks. Consequently, its size and site must be deduced through multiple converging lines of geological and paleontological evidence. Previous and new evidence includes the catastrophically emplaced Alamo Breccia, tsunamites, shock-metamorphosed quartz grains, carbonate accretionary lapilli, an iridium anomaly, sub-Breccia clastic injection, deep-water Breccia channels, and ejecta material. We now demonstrate, on the basis of conodont microfossils in carbonate ejecta clasts within lapillistone blocks and...