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Graptolite faunas exhibited strong biogeographical differentiation during the Early Palaeozoic, particularly in the Ordovician. Skevington recognized two major faunal provinces, the high to mid palaeolatitude ‘Atlantic Province’ and the low-palaeolatitude ‘Pacific Province’. Subsequent workers have generally accepted this pattern of graptolite distribution, but the controls on this pattern have been the subject of considerable debate. Two competing models have emerged: a surface water temperature model and a depth stratification model. It is likely that the some of the physical and chemical oceanic factors that vary with latitude may also vary in a similar way along an onshore to offshore...

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