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Laboratory cultures of several species of benthic foraminifera were grown under controlled physical and chemical conditions during months-long experiments carried out at the University of South Carolina in 2001 and 2002. A dozen experimental culture chambers contained a c. 1–3 mm layer of trace-metal free silica substrate, and were continuously flushed with water from a large (1600 L) seawater reservoir with known, constant temperature and composition (δ18O(water), carbonate system chemistry, and trace element concentrations). Each year, in most of the culture chambers, one or more species reproduced, producing hundreds of juveniles which grew into size classes...

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