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Understanding of biotic response to global change in Earth history is a prime goal in Earth-Life system research (Culver and Rawson, 2000; Lane et al., 2000). The evolution of the Phanerozoic reef ecosystem reflects changes triggered by evolutionary innovations and variations in global and regional controls at different scales. The study of ancient reefs requires bridging across disciplines and timescales and a comparative approach leading to the recognition of trends and patterns. Understanding the ecological change of ancient reefs is also crucial to evaluate threats to modern coral reefs.

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