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An analysis of the lithofacies and trace fossils from a Middle Cambrian deltaic sequence in the Cantabrian Mountains, northwest Spain, indicates the importance of energy conditions, as evidenced by bed thickness and the presence of sand/mud alternations, in determining the distribution of trace fossils. This facies control of trace fossils is particularly strong for trilobite tracks and U-rubes.

An analysis of Arenicolites and Diplocraterion demonstrates that the size of the trace fossil can be taken as an indicator of onshore/offshore conditions.

Thirteen ichnogenera are described, including the new ichnospecies Monomorphichnus pectenensis, Rusophycus ramellen-sis, and Teichichnus ovillus.

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