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The late Miocene-early Pliocene Lake Pannon was a large, long-lived, brackish lake that occupied the Pannonian Basin system in Central Europe. Traditionally, the stratigraphic subdivision of its several-kilometer-thick sediment pile has been based on fossils of endemic mollusks. For a long time, however, stratigraphers were misled by the unconventional architecture of the lacustrine-deltaic sequence. Due to progradation, the increasingly younger sediment packages have been deposited horizontally next to each other instead of forming a purely vertical succession. This pattern was fully recognized recently by seismic exploration of the basin. Newly interpreted biostratigraphy using anagenetically evolving mollusk lineages (in the...

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